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Leathery sea squirt | ![]() |
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Styela clavaHerdman, 1881![]() USGS - http://nas.er.usgs.gov/tunicates/ |
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Taxonomy |
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Description |
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| Styela clava is a large, club-shaped solitary ascidian with a tough leathery body wall (test), growing up to 160mm long. It consists of an elongated, cylindrical body on top of a stalk of variable length. It can be brownish-white, yellowish-brown or reddish-brown. There are two short siphons towards the top of the organism, each with a 4-lobed opening. The body has conspicuous tubercles and rounded swellings on the upper portion and rounded longitudinal ridges on the lower half. The stalk surface is creased. Internally, the gut is a simple U-shaped loop. | |||||||||||||||
Diagram |
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![]() Diagram adapted from Kott, 1985; Christiansen and Thomsen, 1981 |
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Similar Species |
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Styela spp.
MORE (Common Names, Synonyms, Identification Notes, Similar Species, Morphology)... |
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Reproduction and Growth |
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| S. clava is a hermaphrodite but male and female gonads mature at different times, hence they are not self-fertile. Fertilisation is external and eggs and larvae planktonic for between one to three days, after which they settle and metamorphose into the sessile adult. Reproduction occurs throughout all but the coldest periods (when water temperatures <15 degrees C). | |||||||||||||||
Habitat |
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| Present on coasts in low wave energy environments and sheltered embayments in the upper sublittoral zone to at least 25 m depth. As a fouling species, it is common on rocks and pylons and can reach densities of 500-1500 individuals per square metre. It is a hardy species, capable of withstanding salinity changes and temperature fluctuations. | |||||||||||||||
Feeding Preferences |
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Trophic status: Suspension Feeder S. clava feeds on suspended matter such as phytoplankton, zooplankton and organic materials. |
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Predators |
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| Juveniles are eaten by snails such as Mitrella lunata and fish such as Tautogolabrus adspersus. There are no recorded consumers of adult S. clava in northwestern Europe. | |||||||||||||||
Impacts |
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S. clava is a fast growing, suspension feeder that can reach densities of 500-1500 individuals per square metre. As fouling species it can have negative impacts on native and aquaculture species through competition for space and food as well as predation of larvae from the water column. It can also occur as fouling on vessels, aquaculture and fishing equipment and other artificial structures. MORE (Impact information)... |
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Vectors |
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Control Options |
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Click here to enter the Rapid Response Toolbox to search for potential control options. |
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Additional Information |
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Key ReferencesBerman, J., Harris, L., Lambert, W., Buttrick, M., Dufresne, M. (1992). Recent invasions of the Gulf of Maine: three contrasting ecological histories. Conservation and Biology 6:435-441.Christiansen, J. , Thomsen, J. C. (1981). Styela clava Herdman, 1882, a species new to the Danish fauna (Tunicata, Ascidiacea). Steenstrupia 7:15-24. Eno, C.N., Clark, R.A., Sanderson, W.G., (1997). Non-native marine species in British waters: a review and directory. Joint Nature Conservation Committee (JNCC), Peterborough. Holmes, N. (1976). Occurrence of the ascidian Styela clava Herdman in Hobsons Bay, Victoria: a new record for the southern hemisphere. Proceedings of the Royal Society of Victoria 88(1-2):115-116. Kott, P. (1985). The Australian Ascidiacea. Part 1: Phlebobranchia and Stolidobranchia. Memoirs of the Queensland Museum 23:1-438. Lambert, C.C., Lambert, G (1998). Non-indigenous ascidians in southern California harbors and marinas. Marine Biology 130:675-688. Lutzen, J. (1999). Styela clava Herdman (Urochordata, Ascidiacea), a successful immigrant to North-West Europe: ecology, propagation and chronology of spread. Helgolaender Meeresuntersuchungen 52:383-391. Millar, R.H. (1960). The identity of the ascidians Styela mammiculata Carlisle and S. clava Herdman. Journal of the Marine Biological Association of the United Kingdom 39:509-511. Osman, R.W., Whitlatch, R.B. (1999). Ecological interactions of invading ascidians within epifaunal communities of southern New England. IN: Marine bioinvasions: Proceedings of conference January 24-27 1999, (Pederson, J. Eds) Massachusetts Institute of Technology, Cambridge, Massachusetts164-174. | |||||||||||||||
| MORE - References... | |||||||||||||||
| Summary Page Last Modified: | 4/23/2002 |
| Details Pages Last Modified: | 5/11/2004 |
| Distribution Last Modified: | 7/12/2006 |
Please use the following citation for this material
NIMPIS (2002). Styela clava species summary. National Introduced Marine Pest Information System (Eds: Hewitt C.L., Martin R.B., Sliwa C., McEnnulty, F.R., Murphy, N.E., Jones T. & Cooper, S.). Web publication <http://crimp.marine.csiro.au/nimpis>, Date of access: 2/10/2010
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